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It comes from the 2011 animated movie Happy Feet 2. amp.knowyourmeme.com In a side plot, two krill (tiny shrimp-like crustaceans) named Will and Bill (voiced by Brad Pitt and Matt Damon) swim in a massive identical swarm. When Bill remarks that they all look the same, Will replies something like: “Not me, Bill. There’s only one of me in all the world. I am one in a krillion.”“Krillion” is a portmanteau of “krill” + “million” (or similar large-number words), playing on the idea that even though there are enormous numbers of nearly identical krill, Will insists he’s unique.